Action Controller is the C in MVC. After the router has determined which controller to use for a request, the controller is responsible for making sense of the request and producing the output.
The controller will receive the request, fetch or save data from a model and use a view to create output. A controller can be thought of as a middleman between models and views. It makes the model data available to the view so it can display to the user, and it saves or updates user data to the model.